Aegilops species are not native to North America; however,
Ae. triuncialis, Ae. cylindrica and Ae. ovata
survive as adventitious species in the United States. Ae.
triuncialis is the most widely spread Aegilops species in
the world. This species has become a troublesome weed on the
rangelands of California and Pennsylvania (van Slageren 1994).
However, there is little or no awareness about its existence in
Pennsylvania (Hartwig N: pers commun). Ae. cylindrica is also
troublesome in fields and pastures. Its growth on the edges and
within wheat fields is also troublesome. Gene transfer between wheat
(Triticum aestivum) and Ae. cylindrica in the field may
pose a potential problem since Ae. cylindrica may receive
traits, such as herbicide resistance, from transgenic wheat cultivars
through natural introgressive hybridization (Seefeldt et al. 1998;
Zemetra et al. 1998). The geographical distribution of Ae.
cylindrica was surveyed in 1993 by USDA-ARS (http://
www.janr.unl.edu/jgg/maps/). Ae. cylindrica is a
serious weed in Northern Oregon, but not so much problematic in
California. The distribution of Ae. triuncialis in North
America is not well documented. However, information from herbarium
specimen indicates a wide distribution area in the central regions of
California. We had an opportunity to collect 17 populations of
Aegilops species in California and Oregon from 28 June to 6 July,
1999, which is described in this report.
The collection sites and areas covered are presented in Table
1 and Fig. 1.
Aegilops triuncialis (1-6)
In the ranges of south of Milton and San Andreas, California, Ae
triuncialis was found in massive stands. At Winters, however, it
was found in dry and open ground between a dismantled railroad track
and forage-road. Ae. triuncialis is widely distributed at
altitude ranges of 300 m to 1000 m at the Experimental Station,
University of California at Hopland. Ae. triluncialis is
troublesome weed because of its narrow leaves and barbed spikes,
which are not preferred by sheep for grazing. Burning is applied to
reduce population size of Ae. triuncialis at the Experimental
Station. We also found Ae. triuncialis on a southwest facing
road-cut and a slope above the road-cut located three miles northeast
of Nevada City. We did not find Ae. triuncialis in Oregon.
Aegilops cylindrica (7-16)
California was not widely infested with Ae. cylindrica. In
Crestline, we found Ae. cylindrica growing together with wheat
plants at the roadside. At Santa Barbara, Ae. cylindrica
survives at the edge of an old pasture, that used to be a
cattle-feeding lot and now abandoned as grassland. It is thought that
Ae. cylindrica might have spread as a contaminant of
cattle feed. There is herbarium specimen collected in the Santa
Cruz Island, near Santa Barbara (van Slageren 1994). Ae.
cylindrica was found at roadsides and edges of grasslands from
Yreka to Montague in the Siskiyou county. At the east of Montague, it
was found in massive stands in a grassland leading to pastures. There
is a report that Ae. cylindrica was found at Tulelake (Waines
JG: pers commun), but we were unable to find it. In California, the
distribution of Ae. cylindrica was restricted to the roadsides
and edge of pastures.
Wasco, Gillian and Morrow are known to be the most infested Counties
of Ae. cylindrica in Northern Oregon. As summarized in
Table 1, Ae. cyiindrica was found at
the edges of wheat and barley fields and feed lots. At the edge of
wheat field of Condon, we could easily find hybrid plants between
wheat and Ae. cylindrica (Fig. 2).
Hybrid plants were vigorous and their spikelets were sterile. Hybrid
plants were also found in a wheat field at the corner of Ella
Road and Rietniann Lane of lone, Morrow County. In Northern Oregon,
Ae. cylindrica survives at the edge of wheat field and invades
into wheat field where natural hybrids are formed. It means that
Triticum and Aegilops are in the process of
evolution.
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